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Showed that tumors can be caused by viruses
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Discovered the ability of genes to change position on a chromosome
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Proved genetic material is DNA
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One of the first to study chemotherapy for cancer treatments and developed the technique of using human tissue culture rather than laboratory mice to test effects of potential drugs on cancer cells.
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Discovered drugs to treat leukemia and to make organ transplants possible
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Discovered acquired immunity and immune tolerance
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Discovered the structure of DNA
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Discovered the specificity of cell-mediated immunity
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Discovered how cells repair damaged DNA, which can lead to new ways to treat cancer and other diseases
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Helped develop gene targeting, which allows scientist to disable individual genes to understand what they do