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  Discovered the fundamental laws of inheritence. Genes come in pairs-inherited as direct units.
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  Established the chromosomal theory of inheritance.
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  Showed that Fred Grffiths "transforming principle" was DNA.
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  Isolated DNA from different organisms and measured the levels of each of the four nitrogenous bases.
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  Produced X-ray crystallography pictures of BDNA which Watson and Crick use to determine the structure of double-standard DNA.
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  Proved phage DNA, and not protein was the genetic material
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  Solved the structure of DNA
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  Solved the structure of DNA. Also, reasoned that the sequence of nucleotides in the molecule must function as a code. He called this Central Dogma.
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  Made a cell free system that replicates DNA.
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  Showed that rRNA was not the template for building proteins.