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He was born in Russia.
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He married natalia sirvertzera.Natalia was a geneticist and had theodosius daugther named sophie.
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He went to america on a shcolarship to the international Education of the rockefeller foundation.He had pioneered use of fruit flies in a genetic experiment.
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he publishs book like modern synthesis,the synthesis of evolutionary biology with genetics, and genetics of the origins of species.
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In 1937 he won Daneil Giruad Elliot medal.He won it for his work of Genetics and the Origin of Species.
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his wife died of coronary thromebosis.
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He was diagnosed with lukemia and had a few months to live.
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he was the first president of bga.
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he went to calforina for treament.
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he died of heart failure.